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macrophage endocytosis herpetomonas muscarum reservosome cell surface granules trypanosoma cruzi superior colliculus ecto phosphatase trypanosomatid leishmania herpetomonas samuelpessoai vitro cruzi epimastigote neural stem tcrlp squalene synthase estudo inhibitor tcrab localization inorganic phosphate characterization parasite stem cells activity mineral ecto atpase sterol biosynthesis efeito